The domestic dog (Canis lupus familiaris), is a subspecies of the gray wolf (Canis lupus), a member of the Canidae family of the mammilian order Carnivora. The term "domestic dog" is generally used for both domesticated and feral varieties. The dog may have been the first animal to be domesticated, and has been the most widely kept workingSony VAIO PCG-8131M battery, hunting, and companion animal in human history. The word "dog" may also mean the male of a canine species, as opposed to the word "bitch" for the female of the species.
The present lineage of dogs was domesticated from gray wolves about 15,000 years ago.[5] Remains of domesticated dogs have been found in Siberia and Belgium from about 33,000 years ago. None of these early domestication lineages seem to have survived the Last Glacial MaximumSony VAIO PCG-8152M battery. Although mDNA testing suggests a split between dogs and wolves around 100,000 years ago, no specimens prior to 33,000 years ago are clearly morphologically domesticated dog.
Dogs' value to early human hunter-gatherers led to them quickly becoming ubiquitous across world cultures. Dogs perform many roles for people, such as hunting, herding, pulling loads, protection, assisting police and military, companionship, and, more recently, aiding handicapped individualsSony VAIO PCG-31311M battery. This impact on human society has given them the nickname "Man's Best Friend" in the Western world. In some cultures, dogs are also source of meat. In 2001, there were estimated to be 400 million dogs in the world.[11]
Most breeds of dogs are at most a few hundred years old, having been artificially selected for particular morphologies and behaviors by people for specific functional roles. Through this selective breeding, the dog has developed into hundreds of varied breedsSony VAIO PCG-31111M battery, and shows more behavioral and morphological variation than any other land mammal.[12] For example, height measured to the withers ranges from 6 inches (150 mm) in the Chihuahua to about 2.5 feet (0.76 m) in the Irish Wolfhound; color varies from white through grays (usually called "blue") to black, and browns from light (tan) to dark ("red" or "chocolate") in a wide variation of patternsSony VAIO PCG-8112M battery; coats can be short or long, coarse-haired to wool-like, straight, curly, or smooth.[13] It is common for most breeds to shed this coat.
Etymology and related terminology
"Dog" is the common use term that refers to members of the subspecies Canis lupus familiaris (canis, "dog"; lupus, "wolf"; familiaris, "of a household" or "domestic"). The term can also be used to refer to a wider range of related species, such as the members of the genus CanisSony VAIO PCG-7186M battery, or "true dogs", including the wolf, coyote, and jackals, or it can refer to the members of the tribe Canini, which would also include the African wild dog, or it can be used to refer to any member of the family Canidae, which would also include the foxes, bush dog, raccoon dog, and others.[14] Some members of the family have "dog" in their common names, such as the raccoon dog and the African wild dogSony VAIO PCG-7171M battery. A few animals have "dog" in their common names but are not canids, such as the prairie dog.
The English word "dog" comes from Middle English dogge, from Old English docga, a "powerful dog breed".[15] The term may derive from Proto-Germanic *dukkōn, represented in Old English finger-docce ("finger-muscle").Sony VAIO PCG-9Z1M batteryThe word also shows the familiar petname diminutive -ga also seen in frogga "frog", picga "pig", stagga "stag", wicga "beetle, worm", among others.[17] Due to the archaic structure of the word, the term dog may ultimately derive from the earliest layer of Proto-Indo-European vocabulary, reflecting the role of the dog as the earliest domesticated animal. Sony VAIO PCG-5S1M battery
In 14th-century England, "hound" (from Old English: hund) was the general word for all domestic canines, and "dog" referred to a subtype of hound, a group including the mastiff. It is believed this "dog" type of "hound" was so common, it eventually became the prototype of the category “hound”.[19] By the 16th century, dog had become the general wordSony VAIO PCG-5P1M battery, and hound had begun to refer only to types used for hunting.[20] Hound, cognate to German Hund, Dutch hond, common Scandinavian hund, and Icelandic hundur, is ultimately derived from the Proto-Indo-European *kwon- "dog", found in Welsh ci (plural cwn), Latin canis, Greek kýōn, and Lithuanian šuõ.[21]
In breeding circles, a male canine is referred to as a dog, while a female is called a bitch (Middle English biccheSony VAIO PCG-5N2M battery, from Old English bicce, ultimately from Old Norse bikkja). A group of offspring is a litter. The father of a litter is called the sire, and the mother is called the dam. Offspring are, in general, called pups or puppies, from French poupée, until they are about a year old. The process of birth is whelping, from the Old English word hwelp (cf. German Welpe, Dutch welp, Swedish valpa, Icelandic hvelpur) Sony VAIO PCG-3C2M battery.
Taxonomy
In 1753, the father of modern biological taxonomy, Carl Linnaeus, listed among the types of quadrupeds familiar to him, the Latin word for dog, canis. Among the species within this genus, Linnaeus listed the fox, as Canis vulpes, wolves (Canis lupus), and the domestic dog, (Canis canis; see File:Linnaeus - Regnum Animale (1735).png) Sony VAIO PCG-8161M battery.
In later editions, Linnaeus dropped Canis canis and greatly expanded his list of the Canis genus of quadrupeds, and by 1758 included alongside the foxes, wolves, and jackels and many more terms that are now listed as synonyms for domestic dog, including aegyptius (hairless dog), aquaticus, (water dog), and mustelinus (literally “badger dog)” Sony VAIO PCG-8141M battery. Among these were two that later experts have been widely used for domestic dogs as a species: Canis domesticus and, most predominantly, Canis familiaris, the “common” or “familiar” dog.[23]
The domestic dog was accepted as a species in its own right until overwhelming evidence from behavior, vocalizations, morphology, and molecular biology led to the contemporary scientific understanding that a single speciesSony VAIO PCG-3J1M battery, the gray wolf, is the common ancestor for all breeds of domestic dogs. In recognition of this fact, the domestic dog was reclassified in 1993 as Canis lupus familiaris, a subspecies of the gray wolf Canis lupus, by the Smithsonian Institution and the American Society of Mammalogists. C. l. familiaris is listed as the name for the taxon that is broadly used in the scientific community and recommended by ITIS, although Canis familiaris is a recognised synonym. Sony VAIO PCG-3H1M battery
Since that time, C. domesticus and all taxa referring to domestic dogs or subspecies of dog listed by Linnaeus, Johann Friedrich Gmelin in 1792, and Christian Smith in 1839, lost their subspecies status and have been listed as taxonomic synonyms for Canis lupus familiaris
History and evolution
Main articles: Origin of the domestic dog and Gray wolf
Ancient Greek rhyton in the shape of a dog's head, made by Brygos, early 5th century BC. Jérôme Carcopino Museum, Department of Archaeology, AleriaSony VAIO PCG-3F1M battery
Domestic dogs inherited complex behaviors from their wolf ancestors, which would have been pack hunters with complex body language. These sophisticated forms of social cognition and communication may account for their trainability, playfulness, and ability to fit into human households and social situations, and these attributes have given dogs a relationship with humans that has enabled them to become one of the most successful species on the planet today.
Sony VAIO PCG-3C1M battery
Although experts largely disagree over the details of dog domestication, it is agreed that human interaction played a significant role in shaping the subspecies.[29] Domestication may have occurred initially in separate areas particularly Siberia and Europe. Currently it is thought domestication of our current lineage of dog occurred sometime as early as 15,000 years ago and arguably as late as 8500 years agoSony VAIO PCG-9Z2L battery. Shortly after the latest domestication, dogs became ubiquitous in human populations, and spread throughout the world.
Emigrants from Siberia likely crossed the Bering Strait with dogs in their company, and some experts[who?] suggest the use of sled dogs may have been critical to the success of the waves that entered North America roughly 12,000 years ago,[citation needed] although the earliest archaeological evidence of dog-like canids in North America dates from about 9,000 years agoSony VAIO PCG-9Z1L battery.[30] Dogs were an important part of life for the Athabascan population in North America, and were their only domesticated animal. Dogs also carried much of the load in the migration of the Apache and Navajo tribes 1,400 years ago. Use of dogs as pack animals in these cultures often persisted after the introduction of the horse to North America. Sony VAIO PCG-9131L battery
The current consensus among biologists and archaeologists is that the dating of first domestication is indeterminate, although more recent evidence shows isolated domestication events as early as 33,000 years ago. There is conclusive evidence the present lineage of dogs genetically diverged from their wolf ancestors at least 15,000 years agoSony VAIO PCG-8161L battery, but some believe domestication to have occurred earlier.[29] Evidence is accruing that there were previous domestication events, but that those lineages died out.[7]
It is not known whether humans domesticated the wolf as such to initiate dog's divergence from its ancestors, or whether dog's evolutionary path had already taken a different course prior to domesticationSony VAIO PCG-8152L battery. For example, it is hypothesized that some wolves gathered around the campsites of paleolithic camps to scavenge refuse, and associated evolutionary pressure developed that favored those who were less frightened by, and keener in approaching, humans.
Tesem, an old Egyptian sighthound-like dog.
The bulk of the scientific evidence for the evolution of the domestic dog stems from morphological studies of archaeological findings and mitochondrial DNA studiesSony VAIO PCG-8141L battery. The divergence date of roughly 15,000 years ago is based in part on archaeological evidence that demonstrates the domestication of dogs occurred more than 15,000 years ago, and some genetic evidence indicates the domestication of dogs from their wolf ancestors began in the late Upper Paleolithic close to the Pleistocene/Holocene boundary, between 17,000 and 14,000 years ago. Sony VAIO PCG-8131L battery But there is a wide range of other, contradictory findings that make this issue controversial.[citation needed] There are findings beginning currently at 33,000 years ago distinctly placing them as domesticated dogs evidenced not only by shortening of the muzzle but widening as well as crowding of teeth.
Archaeological evidence suggests the latest dogs could have diverged from wolves was roughly 15,000 years ago, although it is possible they diverged much earlier. Sony VAIO PCG-81312L battery In 2008, a team of international scientists released findings from an excavation at Goyet Cave in Belgium declaring a large, toothy canine existed 31,700 years ago and ate a diet of horse, musk ox and reindeer.[37]
Prior to this Belgian discovery, the earliest dog fossils were two large skulls from Russia and a mandible from Germany dated from roughly 14,000 years ago. Sony VAIO PCG-81214L batteryRemains of smaller dogs from Natufian cave deposits in the Middle East, including the earliest burial of a human being with a domestic dog, have been dated to around 10,000 to 12,000 years ago.[5][38] There is a great deal of archaeological evidence for dogs throughout Europe and Asia around this period and through the next two thousand years (roughly 8,000 to 10,000 years ago) Sony VAIO PCG-81115L battery, with fossils uncovered in Germany, the French Alps, and Iraq, and cave paintings in Turkey.[24] The oldest remains of a domesticated dog in the Americas were found in Texas and have been dated to about 9,400 years ago.[39]
DNA studies
DNA studies have provided a wide range of possible divergence dates, from 15,000 to 40,000 years ago,[5] to as much as 100,000 to 140,000 years ago.[40] These results depend on a number of assumptions. Sony VAIO PCG-81114L battery Genetic studies are based on comparisons of genetic diversity between species, and depend on a calibration date. Some estimates of divergence dates from DNA evidence use an estimated wolf-coyote divergence date of roughly 700,000 years ago as a calibration.[41] If this estimate is incorrect, and the actual wolf-coyote divergence is closer to one or two million years ago, or more, then the DNA evidence that supports specific dog-wolf divergence dates would be interpreted very differentlySony VAIO PCG-81113L battery.
Furthermore, it is believed the genetic diversity of wolves has been in decline for the last 200 years, and that the genetic diversity of dogs has been reduced by selective breeding. This could significantly bias DNA analyses to support an earlier divergence date. The genetic evidence for the domestication event occurring in East Asia is also subject to violations of assumptionsSony VAIO PCG-7142L battery. These conclusions are based on the location of maximal genetic divergence, and assume hybridization does not occur, and that breeds remain geographically localized. Although these assumptions hold for many species, there is good reason to believe that they do not hold for canines.[24]
Genetic analyses indicate all dogs are likely descended from a handful of domestication events with a small number of founding females, Sony VAIO PCG-7141L battery although there is evidence domesticated dogs interbred with local populations of wild wolves on several occasions.[5] Data suggest dogs first diverged from wolves in East Asia, and these domesticated dogs then quickly migrated throughout the world, reaching the North American continent around 8000 BC.[5] The oldest groups of dogs, which show the greatest genetic variability and are the most similar to their wolf ancestorsSony VAIO PCG-71111L battery, are primarily Asian and African breeds, including the Basenji, Lhasa Apso, and Siberian Husky.[43] Some breeds thought to be very old, such as the Pharaoh Hound, Ibizan Hound, and Norwegian Elkhound, are now known to have been created more recently.[43]
A great deal of controversy surrounds the evolutionary framework for the domestication of dogs.[24] Although it is widely claimed that "man domesticated the wolf," Sony VAIO PCG-61411L battery man may not have taken such a proactive role in the process.[24] The nature of the interaction between man and wolf that led to domestication is unknown and controversial. At least three early species of the Homo genus began spreading out of Africa roughly 400,000 years ago, and thus lived for a considerable time in contact with canine species. Sony VAIO PCG-61112L battery
Despite this, there is no evidence of any adaptation of canine species to the presence of the close relatives of modern man. If dogs were domesticated, as believed, roughly 15,000 years ago, the event (or events) would have coincided with a large expansion in human territory and the development of agricultureSony VAIO PCG-61111L battery. This has led some biologists to suggest one of the forces that led to the domestication of dogs was a shift in human lifestyle in the form of established human settlements. Permanent settlements would have coincided with a greater amount of disposable food and would have created a barrier between wild and anthropogenic canine populations. Sony VAIO PCG-5T4L battery
Roles with humans
A Siberian Husky used as a pack animal
Early roles
Wolves, and their dog descendants, would have derived significant benefits from living in human camps—more safety, more reliable food, lesser caloric needs, and more chance to breed.[45] They would have benefited from humans’ upright gait that gives them larger range over which to see potential predators and prey, as well as color vision that, at least by day, gives humans better visual discrimination. Sony VAIO PCG-5T3L batteryCamp dogs would also have benefitted from human tool use, as in bringing down larger prey and controlling fire for a range of purposes.[45]
Humans would also have derived enormous benefit from the dogs associated with their camps.[46] For instance, dogs would have improved sanitation by cleaning up food scraps.[46] Dogs may have provided warmth, as referred to in the Australian Aboriginal expression “three dog night” (an exceptionally cold night) Sony VAIO PCG-5T2L battery, and they would have alerted the camp to the presence of predators or strangers, using their acute hearing to provide an early warning.[46]
Anthropologists believe the most significant benefit would have been the use of dogs' sensitive sense of smell to assist with the hunt.[46] The relationship between the presence of a dog and success in the hunt is often mentioned as a primary reason for the domestication of the wolfSony VAIO PCG-5S3L battery, and a 2004 study of hunter groups with and without a dog gives quantitative support to the hypothesis that the benefits of cooperative hunting was an important factor in wolf domestication.[47]
The cohabitation of dogs and humans would have greatly improved the chances of survival for early human groups, and the domestication of dogs may have been one of the key forces that led to human success. Sony VAIO PCG-5S2L battery
A British Bulldog shares a day at the park.
As pets
"The most widespread form of interspecies bonding occurs between humans and dogs"[46] and the keeping of dogs as companions, particularly by elites, has a long history.[49] However, pet dog populations grew significantly after World War II as suburbanization increased.[49] In the 1950s and 1960s, dogs were kept outside more often than they tend to be today Sony VAIO PCG-5S1L battery (using the expression “in the doghouse” to describe exclusion from the group signifies the distance between the doghouse and the home) and were still primarily functional, acting as a guard, children’s playmate, or walking companion. From the 1980s, there have been changes in the role of the pet dog, such as the increased role of dogs in the emotional support of their owners. Sony VAIO PCG-5R2L battery People and dogs have become increasingly integrated and implicated in each other’s lives,[52] to the point where pet dogs actively shape the way a family and home are experienced.[53]
There have been two major trends in the changing status of pet dogs. The first has been the ‘commodification’ of the dog, shaping it to conform to human expectations of personality and behaviour.[53] The second has been the broadening of the concept of the family and the home to include dogs-as-dogs within everyday routines and practices. Sony VAIO PCG-5R1L battery
There are a vast range of commodity forms available to transform a pet dog into an ideal companion.[54] The list of goods, services and places available is enormous: from dog perfumes, couture, furniture and housing, to dog groomers, therapists, trainers and care-takers, dog cafes, spas, parks and beaches, and dog hotels, airlines and cemeteries. Sony VAIO PCG-5P4L battery While dog training as an organized activity can be traced back to the 18th century, in the last decades of the 20th century it became a high profile issue as many normal dog behaviors such as barking, jumping up, digging, rolling in dung, fighting, and urine marking became increasingly incompatible with the new role of a pet dog.[55] Dog training books, classes and television programs proliferated as the process of commodifying the pet dog continued. Sony VAIO PCG-5P2L battery
A pet dog taking part in Christmas traditions
The majority of contemporary dog owners describe their dog as part of the family,[53] although some ambivalence about the relationship is evident in the popular reconceptualization of the dog-human family as a pack.[53] A dominance model of dog-human relationships has been promoted by some dog trainers, such as on the television program Dog WhispererSony VAIO PCG-5N4L battery. However it has been disputed that "trying to achieve status" is characteristic of dog–human interactions.[57] Pet dogs play an active role in family life; for example, a study of conversations in dog-human families showed how family members use the dog as a resource, talking to the dog, or talking through the dog, to mediate their interactions with each other.Sony VAIO PCG-5N2L battery
Another study of dogs’ roles in families showed many dogs have set tasks or routines undertaken as family members, the most common of which was helping with the washing-up by licking the plates in the dishwasher, and bringing in the newspaper from the lawn.[53] Increasingly, human family members are engaging in activities centred on the perceived needs and interests of the dog, or in which the dog is an integral partner, such as Dog Dancing and Doga. Sony VAIO PCG-51513L battery
According to the statistics published by the American Pet Products Manufacturers Association in the National Pet Owner Survey in 2009–2010, it is estimated there are 77.5 million dog owners in the United States.[59] The same survey shows nearly 40% of American households own at least one dog, of which 67% own just one dogSony VAIO PCG-51511L battery, 25% two dogs and nearly 9% more than two dogs. There does not seem to be any gender preference among dogs as pets, as the statistical data reveal an equal number of female and male dog pets. Yet, although several programs are undergoing to promote pet adoption, less than a fifth of the owned dogs come from a shelterSony VAIO PCG-51412L battery.
Work
Dogs have lived and worked with humans in so many roles that they have earned the unique nickname, "man's best friend",[60] a phrase used in other languages as well. They have been bred for herding livestock,[61] hunting (e.g. pointers and hounds),[62] rodent control,[3] guarding, helping fishermen with nets, detection dogs, and pulling loads, in addition to their roles as companionsSony VAIO PCG-51411L battery.
Book of the Hunt, Gaston III, Count of Foix, 1387–88.
Service dogs such as guide dogs, utility dogs, assistance dogs, hearing dogs, and psychological therapy dogs provide assistance to individuals with physical or mental disabilities.[63][64] Some dogs owned by epileptics have been shown to alert their handler when the handler shows signs of an impending seizure, sometimes well in advance of onset, allowing the owner to seek safety, medication, or medical care. Sony VAIO PCG-51312L battery
Dogs included in human activities in terms of helping out humans are usually called working dogs. Dogs of several breeds are considered working dogs. Some working dog breeds include Akita, Alaskan Malamute, Anatolian Shepherd Dog, Bernese Mountain Dog, Black Russian Terrier, Boxer, Bullmastiff, Doberman Pinscher, Dogue de Bordeaux, German Pinscher, German Shepherd, Sony VAIO PCG-51311L batteryGiant Schnauzer, Great Dane, Great Pyrenees, Great Swiss Mountain Dog, Komondor, Kuvasz, Mastiff, Neapolitan Mastiff, Newfoundland, Portuguese Water Dog, Rottweiler, Saint Bernard, Samoyed, Siberian Husky, Standard Schnauzer, and Tibetan Mastiff.
Sports and shows
See also: Conformation show
Owners of dogs often enter them in competitions[67] such as breed conformation shows or sports, including racing and sledding.
In conformation shows, also referred to as breed shows, a judge familiar with the specific dog breed evaluates individual purebred dogs for conformity with their established breed type as described in the breed standardSony VAIO PCG-51211L battery. As the breed standard only deals with the externally observable qualities of the dog (such as appearance, movement, and temperament), separately tested qualities (such as ability or health) are not part of the judging in conformation shows.
As a food source
See also: Dog meat
Fried dog meat offered as speciality, among lamb and fish. Shaoyang, Hunan Province, China
Dog meat is consumed in some East Asian countries, including Korea, China, and Vietnam, a practice that dates back to antiquity. Sony VAIO PCG-41112L battery It is estimated that 13–16 million dogs are killed and consumed in Asia every year.[69] The BBC claims that, in 1999, more than 6,000 restaurants served soups made from dog meat in South Korea.[70] In Korea, the primary dog breed raised for meat, the nureongi (누렁이), differs from those breeds raised for pets that Koreans may keep in their homes. Sony VAIO PCG-3A4L battery
The most popular Korean dog dish is gaejang-guk (also called bosintang), a spicy stew meant to balance the body's heat during the summer months; followers of the custom claim this is done to ensure good health by balancing one's gi, or vital energy of the body. A 19th century version of gaejang-guk explains that the dish is prepared by boiling dog meat with scallions and chili powderSony VAIO PCG-3A3L battery. Variations of the dish contain chicken and bamboo shoots. While the dishes are still popular in Korea with a segment of the population, dog is not as widely consumed as beef, chicken, and pork.
Other cultures, such as Polynesia and pre-Columbian Mexico, also consumed dog meat in their history. However, Western, South Asian, African, and Middle Eastern cultures, in general, regard consumption of dog meat as tabooSony VAIO PCG-3A2L battery. In some places, however, such as in rural areas of Poland, dog fat is believed to have medicinal properties—being good for the lungs for instance.[73]
A CNN report in China dated March 2010 interviews a dog meat vendor who states that most of the dogs that are available for selling to restaurant are raised in special farms but that there is always a chance that a sold dog is someone's lost pet, although dog pet breeds are not considered edible. Sony VAIO PCG-3A1L battery
Health risks to humans
Main article: Dog attack
It is estimated that two percent of the US population, 4.7 million people, are bitten each year.[75] In the 1980s and 1990s the US averaged 17 fatalities per year, while in the 2000s this has increased to 26. 77% of dog bites are from the pet of family or friends, and 50% of attacks occur on the dog owner's property. Sony VAIO PCG-394L battery
A Colorado study found bites in children were less severe than bites in adults.[77] The incidence of dog bites in the U.S. is 12.9 per 10,000 inhabitants, but for boys aged 5 to 9, the incidence rate is 60.7 per 10,000. Moreover, children have a much higher chance to be bitten in the face or neck.[78] Sharp claws with powerful muscles behind them can lacerate flesh in a scratch that can lead to serious infections. Sony VAIO PCG-393L battery
In the UK between 2003 and 2004, there were 5,868 dog attacks on humans, resulting in 5,770 working days lost in sick leave.[80]
In the United States, cats and dogs are a factor in more than 86,000 falls each year.[81] It has been estimated around 2% of dog-related injuries treated in UK hospitals are domestic accidents. The same study found that while dog involvement in road traffic accidents was difficult to quantify, dog-associated road accidents involving injury more commonly involved two-wheeled vehicles. Sony VAIO PCG-391L battery
Toxocara canis (dog roundworm) eggs in dog feces can cause toxocariasis. In the United States, about 10,000 cases of Toxocara infection are reported in humans each year, and almost 14% of the U.S. population is infected.[83] In Great Britain, 24% of soil samples taken from public parks contained T. canis eggs.[84] Untreated toxocariasis can cause retinal damage and decreased vision. Sony VAIO PCG-384L battery Dog feces can also contain hookworms that cause cutaneous larva migrans in humans.
Health benefits for humans
A human cuddles a Doberman puppy.
A growing body of research indicates the companionship of a dog can enhance human physical health and psychological wellbeing.[89] Dog and cat owners have been shown to have better mental and physical health than nonowners, making fewer visits to the doctor and being less likely to be on medication than nonowners. Sony VAIO PCG-383L battery In one study, new pet owners reported a highly significant reduction in minor health problems during the first month following pet acquisition, and this effect was sustained in dog owners through to the end of the study.[91]
In addition, dog owners took considerably more physical exercise than cat owners and people without pets. The group without pets exhibited no statistically significant changes in health or behaviourSony VAIO PCG-382L battery. The results provide evidence that pet acquisition may have positive effects on human health and behaviour, and that for dog owners these effects are relatively long term.[91] Pet ownership has also been associated with increased coronary artery disease survival, with dog owners being significantly less likely to die within one year of an acute myocardial infarction than those who did not own dogs.Sony VAIO PCG-381L battery
Gunnar Kaasen and Balto, the lead dog on the last relay team of the 1925 serum run to Nome.
The health benefits of dogs can result from contact with dogs, not just from dog ownership. For example, when in the presence of a pet dog, people show reductions in cardiovascular, behavioral, and psychological indicators of anxiety. Sony VAIO PCG-7185L battery The benefits of contact with a dog also include social support, as dogs are able to not only provide companionship and social support themselves, but also to act as facilitators of social interactions between humans.[94] One study indicated that wheelchair users experience more positive social interactions with strangers when they are accompanied by a dog than when they are not. Sony VAIO PCG-7184L battery
The practice of using dogs and other animals as a part of therapy dates back to the late 18th century, when animals were introduced into mental institutions to help socialize patients with mental disorders.[96] Animal-assisted intervention research has shown that animal-assisted therapy with a dog can increase a person with Alzheimer’s disease’s social behavioursSony VAIO PCG-7183L battery, such as smiling and laughing.[97] One study demonstrated that children with ADHD and conduct disorders who participated in an education program with dogs and other animals showed increased attendance, increased knowledge and skill objectives, and decreased antisocial and violent behavior compared to those who were not in an animal-assisted program. Sony VAIO PCG-7182L battery
Shelters
Main article: Animal shelter
Every year, between 6 and 8 million dogs and cats enter US animal shelters.[99] The Humane Society of the United States (HSUS) estimates that approximately 3 to 4 million dogs and cats are euthanized yearly in shelters across the United States. However, the percentage of dogs in US animal shelters that are eventually adopted and removed from the sheltersSony VAIO PCG-7181L battery by their new owners has increased since the mid 1990s from around 25% up to around 60–75% in the mid first decade of the 21st century.[101]
Pets entering the shelters are euthanized in countries all over the world because of the lack of financial provisions to take care of these animals. Most shelters complain of not having enough resources to feed the pets and by being constrained to kill them, as the likelihood for all of them to find an owner is very small. In poor countries, euthanasia is usually violentSony VAIO PCG-7174L battery.
Biology
Main article: Dog anatomy
Lateral view of a dog's bone structure.
Domestic dogs have been selectively bred for millennia for various behaviors, sensory capabilities, and physical attributes.[3] Modern dog breeds show more variation in size, appearance, and behavior than any other domestic animal. Nevertheless, their morphology is based on that of their wild ancestors, gray wolves.[3] Dogs are predators and scavengersSony VAIO PCG-7173L battery, and like many other predatory mammals, the dog has powerful muscles, fused wrist bones, a cardiovascular system that supports both sprinting and endurance, and teeth for catching and tearing.
Dogs are highly variable in height and weight. The smallest known adult dog was a Yorkshire Terrier, that stood only 6.3 centimetres (2.5 in) at the shoulder, 9.5 cm (3.7 in) in length along the head-and-body, and weighed only 113 grams (4.0 oz)Sony VAIO PCG-7172L battery. The largest known dog was an English Mastiff which weighed 155.6 kilograms (343 lb) and was 250 cm (98 in) from the snout to the tail.[102] The tallest dog is a Great Dane that stands 106.7 cm (42.0 in) at the shoulder.
Senses
Vision
Dog's visual colour perception compared with humans.
Like most mammals, dogs are dichromats and have color vision equivalent to red-green color blindness in humans (deuteranopia). Dogs are less sensitive to differences in grey shades than humans and also can detect brightness at about half the accuracy of humans. Sony VAIO PCG-7171L battery
The dog's visual system has evolved to aid proficient hunting.[104] While a dog's visual acuity is poor (that of a poodle's has been estimated to translate to a Snellen rating of 20/75[104]), their visual discrimination for moving objects is very high; dogs have been shown to be able to discriminate between humans (e.g., identifying their owner) at a range of between 800 and 900 m, however this range decreases to 500–600 m if the object is stationary. Sony VAIO PCG-7162L battery
Dogs have a temporal resolution of between 60 and 70 Hz, which explains why many dogs struggle to watch television, as most such modern screens are optimized for humans at 50–60 Hz.[108] Dogs can detect a change in movement that exists in a single diopter of space within their eye. Humans, by comparison, require a change of between 10 and 20 diopters to detect movementSony VAIO PCG-7161L battery.
The physiology of a dog ear.
As crepuscular hunters, dogs often rely on their vision in low light situations: They have very large pupils, a high density of rods in the fovea, an increased flicker rate, and a tapetum lucidum.[104] The tapetum is a reflective surface behind the retina that reflects light to give the photoreceptors a second chance to catch the photons. There is also a relationship between body size and overall diameter of the eyeSony VAIO PCG-7154L battery. A range of 9.5 and 11.6 mm can be found between various breeds of dogs. This 20% variance can be substantial and is associated as an adaptation toward superior night vision.[111]
The eyes of different breeds of dogs have different shapes, dimensions, and retina configurations.[112] Many long-nosed breeds have a "visual streak" – a wide foveal region that runs across the width of the retina and gives them a very wide field of excellent visionSony VAIO PCG-7153L battery. Some long-muzzled breeds, in particular, the sighthounds, have a field of vision up to 270° (compared to 180° for humans). Short-nosed breeds, on the other hand, have an "area centralis": a central patch with up to three times the density of nerve endings as the visual streak, giving them detailed sight much more like a human's. Some broad-headed breeds with short noses have a field of vision similar to that of humans. Sony VAIO PCG-7152L battery
Most breeds have good vision, but some show a genetic predisposition for myopia – such as Rottweilers, with which one out of every two has been found to be myopic.[104] Dogs also have a greater divergence of the eye axis than humans, enabling them to rotate their pupils farther in any direction. The divergence of the eye axis of dogs ranges from 12–25° depending on the breed.Sony VAIO PCG-7151L battery
Experimentation has proven that dogs can distinguish between complex visual images such as that of a cube or a prism. Dogs also show attraction to static visual images such as the silhouette of a dog on a screen, their own reflections, or videos of dogs; however, their interest declines sharply once they are unable to make social contact with the image.
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Hearing
The frequency range of dog hearing is approximately 40 Hz to 60,000 Hz,[114] which means that dogs can detect sounds far beyond the upper limit of the human auditory spectrum. In addition, dogs have ear mobility, which allows them to rapidly pinpoint the exact location of a sound. Eighteen or more muscles can tilt, rotate, raise, or lower a dog's earSony VPCW21M2E/WI battery. A dog can identify a sound's location much faster than a human can, as well as hear sounds at four times the distance.[116]
Smell
The wet, textured nose of a dog
While the human brain is dominated by a large visual cortex, the dog brain is dominated by an olfactory cortex.[104] The olfactory bulb in dogs is roughly forty times bigger than the olfactory bulb in humans, relative to total brain size, with 125 to 220 million smell-sensitive receptors.[104] The bloodhound exceeds this standard with nearly 300 million receptors. Sony VPCW21C7E battery
Consequently, it has been estimated that dogs, in general, have an olfactory sense ranging from one hundred thousand to one million times more sensitive than a human's. In some dog breeds, such as bloodhounds, the olfactory sense may be up to 100 million times greater than a human's.[117] The wet nose is essential for determining the direction of the air current containing the smellSony VPCW12S1E/WZ battery. Cold receptors in the skin are sensitive to the cooling of the skin by evaporation of the moisture by air currents.[118]
Physical characteristics
Main article: Dog anatomy
Coat
Main article: Coat (dog)
A heavy winter coat with countershading in a mixed-breed dog
The coats of domestic dogs are of two varieties: "double" being common with dogs (as well as wolves) originating from colder climates, made up of a coarse guard hair and a soft down hair, or "single", with the topcoat onlySony VPCW12S1E/W battery.
Domestic dogs often display the remnants of countershading, a common natural camouflage pattern. A countershaded animal will have dark coloring on its upper surfaces and light coloring below,[119] which reduces its general visibility. Thus, many breeds will have an occasional "blaze", stripe, or "star" of white fur on their chest or underside. Sony VPCW12S1E/T battery
Tail
See also: Docking
There are many different shapes for dog tails: straight, straight up, sickle, curled, or cork-screw. As with many canids, one of the primary functions of a dog's tail is to communicate their emotional state, which can be important in getting along with others. In some hunting dogs, however, the tail is traditionally docked to avoid injuries.[121] In some breeds, puppies can be born with a short tail or no tail at allSony VPCW12S1E/P battery.
Types and breeds
Main article: Dog breed
Further information: Dog type
Cavalier King Charles Spaniels demonstrate with-breed variation.
While all dogs are genetically very similar,[5] natural selection and selective breeding have reinforced certain characteristics in certain populations of dogs, giving rise to dog types and dog breeds. Dog types are broad categories based on function, genetics, or characteristics.[123] Dog breeds are groups of animals that possess a set of inherited characteristics that distinguishes them from other animals within the same speciesSony VPCW11S1E/W battery. Modern dog breeds are non-scientific classifications of dogs kept by modern kennel clubs.
Purebred dogs of one breed are genetically distinguishable from purebred dogs of other breeds,[43] but the means by which kennel clubs classify dogs is unsystematic. Systematic analyses of the dog genome has revealed only four major types of dogs that can be said to be statistically distinct.[43] These include the "old world dogs" (e.g., Malamute and Shar Pei) Sony VPCW11S1E/T battery, "Mastiff"-type (e.g., English Mastiff), "herding"-type (e.g., Border Collie), and "all others" (also called "modern"- or "hunting"-type).
Health
Main article: Dog health
Further information: Category:Dog health
See also: CVBD
Dogs are susceptible to various diseases, ailments, and poisons, some of which can affect humans. To defend against many common diseases, dogs are often vaccinated.
A mixed-breed dog
Some breeds of dogs are prone to certain genetic ailments such as elbow or hip dysplasia, blindness, deafness, pulmonic stenosis, cleft palate, and trick knees. Two serious medical conditions particularly affecting dogs are pyometra, affecting unspayed females of all types and agesSony VPCW11S1E/P battery, and bloat, which affects the larger breeds or deep-chested dogs. Both of these are acute conditions, and can kill rapidly. Dogs are also susceptible to parasites such as fleas, ticks, and mites, as well as hookworm, tapeworm, roundworm, and heartworm.
Dogs are highly susceptible to theobromine poisoning, typically from ingestion of chocolate. Theobromine is toxic to dogs because, although the dog's metabolism is capable of breaking down the chemicalSony VPCYA1S1E/B battery, the process is so slow that even small amounts of chocolate can be fatal, especially dark chocolate.
Dogs are also vulnerable to some of the same health conditions as humans, including diabetes, dental and heart disease, epilepsy, cancer, hypothyroidism, and arthritis.[125]
Mortality
Main article: Aging in dogs
The typical lifespan of dogs varies widely among breeds, but for most the median longevity, the age at which half the dogs in a population have died and half are still alive, ranges from 10 to 13 years. Individual dogs may live well beyond the median of their breedSony VPCYA1V9E/B battery.
The breed with the shortest lifespan (among breeds for which there is a questionnaire survey with a reasonable sample size) is the Dogue de Bordeaux, with a median longevity of about 5.2 years, but several breeds, including Miniature Bull Terriers, Bloodhounds, and Irish Wolfhounds are nearly as short-lived, with median longevities of 6 to 7 years. Sony VPCY21S1E/SI battery
The longest-lived breeds, including Toy Poodles, Japanese Spitz, Border Terriers, and Tibetan Spaniels, have median longevities of 14 to 15 years.[129] The median longevity of mixed-breed dogs, taken as an average of all sizes, is one or more years longer than that of purebred dogs when all breeds are averaged. The dog widely reported to be the longest-lived is "Bluey," Sony VPCY21S1E/L batterywho died in 1939 and was claimed to be 29.5 years old at the time of his death; however, the Bluey record is anecdotal and unverified.[131] On December 5, 2011, Pusuke, the world's oldest living dog recognized by Guinness Book of World Records, died aged 26 years and 9 months.[132]
Predation
Although wild dogs, like wolves, are apex predators, they can be killed in territory disputes with wild animals. Sony VPCY21S1E/G batteryFurthermore, in areas where both dogs and other large predators live, dogs can be a major food source for big cats or canines. Reports from Croatia indicate wolves kill dogs more frequently than they kill sheep. Wolves in Russia apparently limit feral dog populations. In Wisconsin, more compensation has been paid for dog losses than livestock.[133] Some wolf pairs have been reported to prey on dogs by having one wolf lure the dog out into heavy brush where the second animal waits in ambush. Sony VPCY11S1E/S battery In some instances, wolves have displayed an uncharacteristic fearlessness of humans and buildings when attacking dogs, to the extent that they[which?] have to be beaten off or killed.[135]
Coyotes and big cats have also been known to attack dogs. Leopards in particular are known to have a predilection for dogs, and have been recorded to kill and consume them regardless of the dog's size or ferocity.[136] Tigers in Manchuria, Indochina, Indonesia, and Malaysia are reputed to kill dogs with the same vigor as leopards. Sony VPCY11S1E battery Striped Hyenas are major predators of village dogs in Turkmenistan, India, and the Caucasus.[138] Reptiles such as alligators and pythons have been known to kill and eat dogs.
Diet
Dog food
Golden Retriever gnawing a pig's foot
Despite their descent from wolves and classification as Carnivora, dogs are variously described in scholarly and other writings as carnivores or omnivores. Unlike obligate carnivores, such as the cat family with its shorter small intestineSony VPCZ11X9E/B battery, dogs can adapt to a wide-ranging diet, and are not dependent on meat-specific protein nor a very high level of protein in order to fulfill their basic dietary requirements. Dogs will healthily digest a variety of foods, including vegetables and grains, and can consume a large proportion of these in their diet. Sony VPCZ11Z9E/B battery
A number of common human foods and household ingestibles are toxic to dogs, including chocolate solids (theobromine poisoning), onion and garlic (thiosulphate, sulfoxide or disulfide poisoning),[144] grapes and raisins, macadamia nuts, as well as various plants and other potentially ingested materialsSony VPCZ12M9E/B battery.
Reproduction
Two dogs copulating on a beach
Main article: Canine reproduction
In domestic dogs, sexual maturity begins to happen around age six to twelve months for both males and females,[3][147] although this can be delayed until up to two years old for some large breeds. This is the time at which female dogs will have their first estrous cycle. They will experience subsequent estrous cycles biannually, during which the body prepares for pregnancySony VPCZ12X9E/X battery. At the peak of the cycle, females will come into estrus, being mentally and physically receptive to copulation.[3] Because the ova survive and are capable of being fertilized for a week after ovulation, it is possible for a female to mate with more than one male.[3]
Dogs bear their litters roughly 56 to 72 days after fertilization,[3][148] with an average of 63 days, although the length of gestation can vary. An average litter consists of about six puppies, Sony VPCZ12Z9E/X battery though this number may vary widely based on the breed of dog. In general, toy dogs produce from one to four puppies in each litter, while much larger breeds may average as many as twelve.
Some dog breeds have acquired traits through selective breeding that interfere with reproduction. Male French Bulldogs, for instance, are incapable of mounting the female. For many dogs of this breed, the female must be artificially inseminated in order to reproduce. Sony VPCZ138GA battery
Neutering
The examples and perspective in this article deal primarily with the United States and do not represent a worldwide view of the subject. Please improve this article and discuss the issue on the talk page. (March 2012)
A feral dog from Sri Lanka nursing her four puppies
Neutering refers to the sterilization of animals, usually by removal of the male's testicles or the female's ovaries and uterus, in order to eliminate the ability to procreate and reduce sex driveSony VPCZ13M9E/B battery. Because of the overpopulation of dogs in some countries, many animal control agencies, such as the American Society for the Prevention of Cruelty to Animals (ASPCA), advise that dogs not intended for further breeding should be neutered, so that they do not have undesired puppies that may have to later be euthanized.[151]
According to the Humane Society of the United States, 3–4 million dogs and cats are put down each year in the United States and many more are confined to cages in shelters because there are many more animals than there are homesSony VPCZ13M9E/X battery. Spaying or castrating dogs helps keep overpopulation down.[152] Local humane societies, SPCAs, and other animal protection organizations urge people to neuter their pets and to adopt animals from shelters instead of purchasing them.
Neutering reduces problems caused by hypersexuality, especially in male dogs.[153] Spayed female dogs are less likely to develop some forms of cancer, affecting mammary glands, ovaries, and other reproductive organs. Sony VPCZ13V9E batteryHowever, neutering increases the risk of urinary incontinence in female dogs,[155] and prostate cancer in males,[156] as well as osteosarcoma, hemangiosarcoma, cruciate ligament rupture, obesity, and diabetes mellitus in either gender.[157]
Intelligence and behavior
Intelligence
Main article: Dog intelligence
The Border Collie is considered to be one of the most intelligent breeds.
The domestic dog has a predisposition to exhibit a social intelligence that is uncommon in the animal world.[104] Dogs are capable of learning in a number of ways, such as through simple reinforcement (e.g., classical or operant conditioning) and by observation. Sony VPCZ13V9E/X battery
Dogs go through a series of stages of cognitive development. As with humans, the understanding that objects not being actively perceived still remain in existence (called object permanence) is not present at birth. It develops as the young dog learns to interact intentionally with objects around it, at roughly 8 weeks of age. Sony VPCZ13Z9E/X battery
Puppies learn behaviors quickly by following examples set by experienced dogs.[104] This form of intelligence is not peculiar to those tasks dogs have been bred to perform, but can be generalized to myriad abstract problems. For example, Dachshund puppies that watched an experienced dog pull a cart by tugging on an attached piece of ribbon in order to get a reward from inside the cart learned the task fifteen times faster than those left to solve the problem on their own. Sony VPCZ21M9E battery
Dogs can also learn by mimicking human behaviors. In one study, puppies were presented with a box, and shown that, when a handler pressed a lever, a ball would roll out of the box. The handler then allowed the puppy to play with the ball, making it an intrinsic reward. The pups were then allowed to interact with the box. Roughly three-quarters of the puppies subsequently touched the leverSony VPCZ21Q9E battery, and over half successfully released the ball, compared to only 6% in a control group that did not watch the human manipulate the lever.[159] Another study found that handing an object between experimenters who then used the object's name in a sentence successfully taught an observing dog each object's name, allowing the dog to subsequently retrieve the item. Sony VPCZ21V9E battery
Sergeant Stubby wearing his uniform and medals. Stubby participated in four offensives and 17 battles.
Dogs also demonstrate sophisticated social cognition by associating behavioral cues with abstract meanings.[104] One such class of social cognition involves the understanding that others are conscious agents. Research has shown that dogs are capable of interpreting subtle social cuesSony VPCEH3T9E battery, and appear to recognize when a human or dog's attention is focused on them. To test this, researchers devised a task in which a reward was hidden underneath one of two buckets. The experimenter then attempted to communicate with the dog to indicate the location of the reward by using a wide range of signals: tapping the bucket, pointing to the bucket, nodding to the bucket, or simply looking at the bucket. Sony VPCEH3N6E battery The results showed that domestic dogs were better than chimpanzees, wolves, and human infants at this task, and even young puppies with limited exposure to humans performed well.[104]
Psychology research has shown that humans´ gaze instinctively moves to the left in order to watch the right side of a person's face, which is related to use of right hemisphere brain for facial recognition, including human facial emotions. Research at the University of Lincoln (2008) shows that dogs share this instinct when meeting a human beingSony VPCEH3N1E battery, and only when meeting a human being (i.e., not other animals or other dogs). As such they are the only non-primate species known to do so.[162][163]
Stanley Coren, an expert on dog psychology, states that these results demonstrated the social cognition of dogs can exceed that of even our closest genetic relatives, and that this capacity is a recent genetic acquisition that distinguishes the dog from its ancestor, the wolf. Sony VPCEH3D0E battery Studies have also investigated whether dogs engaged in partnered play change their behavior depending on the attention-state of their partner.[164] Those studies showed that play signals were only sent when the dog was holding the attention of its partner. If the partner was distracted, the dog instead engaged in attention-getting behavior before sending a play signal. Sony VPCEH3B1E battery
Coren has also argued that dogs demonstrate a sophisticated theory of mind by engaging in deception, which he supports with a number of anecdotes, including one example wherein a dog hid a stolen treat by sitting on it until the rightful owner of the treat left the room.[104] Although this could have been accidental, Coren suggests that the thief understood that the treat's owner would be unable to find the treat if it were out of viewSony VPCEH2Z1E battery. Together, the empirical data and anecdotal evidence points to dogs possessing at least a limited form of theory of mind.[104][164]
A study found a third of dogs suffered from anxiety when separated from others.[165]
A Border Collie named Chaser has learned the names for 1,022 toys after three years of training, so many that her trainers have had to mark the names of the objects lest they forget themselves. This is higher than Rico, another border collie who could remember at least 200 objects. Sony VPCEH2S9E battery
Behavior
Main article: Dog behavior
Further information: Category:Dog training and behavior
Although dogs have been the subject of a great deal of behaviorist psychology (e.g. Pavlov's dog), they do not enter the world with a psychological "blank slate".[104] Rather, dog behavior is affected by genetic factors as well as environmental factors.[104] Domestic dogs exhibit a number of behaviors and predispositions that were inherited from wolves. Sony VPCEH2Q1E battery
The Gray Wolf is a social animal that has evolved a sophisticated means of communication and social structure. The domestic dog has inherited some of these predispositions, but many of the salient characteristics in dog behavior have been largely shaped by selective breeding by humans. Thus some of these characteristics, such as the dog's highly developed social cognition, are found only in primitive forms in grey wolves. Sony VPCEH2P0E battery
The existence and nature of personality traits in dogs have been studied (15329 dogs of 164 different breeds) and five consistent and stable "narrow traits" identified, described as playfulness, curiosity/fearlessness, chase-proneness, sociability and aggressiveness. A further higher order axis for shyness–boldness was also identified. Sony VPCEH2N1E battery
Sleep
Further information: Sleep (non-human)
The average sleep time of a dog is said to be 10.1 hours per day.[169] Like humans, dogs have two main types of sleep: Slow-wave sleep, then Rapid eye movement sleep sleep, the state in which dreams occur.[170]
Dog growl
A new study in Budapest, Hungary has found that dogs are able to tell how big another dog is just by listening to its growl. A specific growl is used by dogs to protect their food. The research also shows that dogs do not lie about their sizeSony VPCEH2M9E battery, and this is the first time research has shown animals can determine another’s size by the sound it makes. The test, using images of many kinds of dogs, showed a small and big dog and played a growl. The result, showed that 20 of the 24 test dogs looked at the image of the appropriate-sized dog first and looked at it longest. Sony VPCEH2M1E battery
Differences from wolves
Some dogs, like this Tamaskan, look very much like wolves.
Physical characteristics
Further information: Wolf
Compared to equally sized wolves, dogs tend to have 20% smaller skulls, 30% smaller brains,[172] as well as proportionately smaller teeth than other canid species.[173] Dogs require fewer calories to function than wolves. It is thought by certain experts that the dog's limp ears are a result of atrophy of the jaw muscles.[173] The skin of domestic dogs tends to be thicker than that of wolvesSony VPCEH2L9E battery, with some Inuit tribes favoring the former for use as clothing due to its greater resistance to wear and tear in harsh weather.[173]
Behavior
Dogs tend to be poorer than wolves at observational learning, being more responsive to instrumental conditioning.[173] Feral dogs show little of the complex social structure or dominance hierarchy present in wolf packs. For example, unlike wolves, the dominant alpha pairs of a feral dog pack do not force the other members to wait for their turn on a meal when scavenging off a dead ungulate as the whole family is free to join inSony VPCEH2J1E battery. For dogs, other members of their kind are of no help in locating food items, and are more like competitors.[173]
Feral dogs are primarily scavengers, with studies showing that unlike their wild cousins, they are poor ungulate hunters, having little impact on wildlife populations where they are sympatric. However, feral dogs have been reported to be effective hunters of reptiles in the Galápagos Islands, Sony VPCEH2H1E battery and free ranging pet dogs are more prone to predatory behavior toward wild animals.
Domestic dogs can be monogamous.[175] Breeding in feral packs can be, but does not have to be restricted to a dominant alpha pair (such things also occur in wolf packs).[176] Male dogs are unusual among canids by the fact that they mostly seem to play no role in raising their puppies, and do not kill the young of other females to increase their own reproductive success. Sony VPCEH2F1E battery Some sources say that dogs differ from wolves and most other large canid species by the fact that they do not regurgitate food for their young, nor the young of other dogs in the same territory.[173]
An Australian Shepherd-Beagle mix displaying mastery of the "sit" command
However, this difference was not observed in all domestic dogs. Regurgitating of food by the females for the young as well as care for the young by the males has been observed in domestic dogsSony VPCEH2E0E battery, dingos as well as in other feral or semi-feral dogs. Regurgitating of food by the females and direct choosing of only one mate has been observed even in those semi-feral dogs of direct domestic dog ancestry. Also regurgitating of food by males has been observed in free-ranging domestic dogs.
Trainability
Dogs display much greater tractability than tame wolves, and are, in general, much more responsive to coercive techniques involving fearSony VPCEH2D0E battery, aversive stimuli, and force than wolves, which are most responsive toward positive conditioning and rewards.[178] Unlike tame wolves, dogs tend to respond more to voice than hand signals.[179]
Mythology
In mythology, dogs often serve as pets or as watchdogs.[180]
In Greek mythology, Cerberus is a three-headed watchdog who guards the gates of Hades.[180] In Norse mythology, a bloody, four-eyed dog called Garmr guards Helheim.[180] In Persian mythology, two four-eyed dogs guard the Chinvat Bridge. Sony VPCEH2C0E batteryIn Philippine mythology, Kimat who is the pet of Tadaklan, god of thunder, is responsible for lightning. In Welsh mythology, Annwn is guarded by Cŵn Annwn[180]
In Judaism and Islam, dogs are viewed as unclean scavengers.[180] In Christianity, dogs represent faithfulness.[180] In Asian countries such as China, Korea, and Japan, dogs are viewed as kind protectorsSony VPCEH1Z1E battery.
The present lineage of dogs was domesticated from gray wolves about 15,000 years ago.[5] Remains of domesticated dogs have been found in Siberia and Belgium from about 33,000 years ago. None of these early domestication lineages seem to have survived the Last Glacial MaximumSony VAIO PCG-8152M battery. Although mDNA testing suggests a split between dogs and wolves around 100,000 years ago, no specimens prior to 33,000 years ago are clearly morphologically domesticated dog.
Dogs' value to early human hunter-gatherers led to them quickly becoming ubiquitous across world cultures. Dogs perform many roles for people, such as hunting, herding, pulling loads, protection, assisting police and military, companionship, and, more recently, aiding handicapped individualsSony VAIO PCG-31311M battery. This impact on human society has given them the nickname "Man's Best Friend" in the Western world. In some cultures, dogs are also source of meat. In 2001, there were estimated to be 400 million dogs in the world.[11]
Most breeds of dogs are at most a few hundred years old, having been artificially selected for particular morphologies and behaviors by people for specific functional roles. Through this selective breeding, the dog has developed into hundreds of varied breedsSony VAIO PCG-31111M battery, and shows more behavioral and morphological variation than any other land mammal.[12] For example, height measured to the withers ranges from 6 inches (150 mm) in the Chihuahua to about 2.5 feet (0.76 m) in the Irish Wolfhound; color varies from white through grays (usually called "blue") to black, and browns from light (tan) to dark ("red" or "chocolate") in a wide variation of patternsSony VAIO PCG-8112M battery; coats can be short or long, coarse-haired to wool-like, straight, curly, or smooth.[13] It is common for most breeds to shed this coat.
Etymology and related terminology
"Dog" is the common use term that refers to members of the subspecies Canis lupus familiaris (canis, "dog"; lupus, "wolf"; familiaris, "of a household" or "domestic"). The term can also be used to refer to a wider range of related species, such as the members of the genus CanisSony VAIO PCG-7186M battery, or "true dogs", including the wolf, coyote, and jackals, or it can refer to the members of the tribe Canini, which would also include the African wild dog, or it can be used to refer to any member of the family Canidae, which would also include the foxes, bush dog, raccoon dog, and others.[14] Some members of the family have "dog" in their common names, such as the raccoon dog and the African wild dogSony VAIO PCG-7171M battery. A few animals have "dog" in their common names but are not canids, such as the prairie dog.
The English word "dog" comes from Middle English dogge, from Old English docga, a "powerful dog breed".[15] The term may derive from Proto-Germanic *dukkōn, represented in Old English finger-docce ("finger-muscle").Sony VAIO PCG-9Z1M batteryThe word also shows the familiar petname diminutive -ga also seen in frogga "frog", picga "pig", stagga "stag", wicga "beetle, worm", among others.[17] Due to the archaic structure of the word, the term dog may ultimately derive from the earliest layer of Proto-Indo-European vocabulary, reflecting the role of the dog as the earliest domesticated animal. Sony VAIO PCG-5S1M battery
In 14th-century England, "hound" (from Old English: hund) was the general word for all domestic canines, and "dog" referred to a subtype of hound, a group including the mastiff. It is believed this "dog" type of "hound" was so common, it eventually became the prototype of the category “hound”.[19] By the 16th century, dog had become the general wordSony VAIO PCG-5P1M battery, and hound had begun to refer only to types used for hunting.[20] Hound, cognate to German Hund, Dutch hond, common Scandinavian hund, and Icelandic hundur, is ultimately derived from the Proto-Indo-European *kwon- "dog", found in Welsh ci (plural cwn), Latin canis, Greek kýōn, and Lithuanian šuõ.[21]
In breeding circles, a male canine is referred to as a dog, while a female is called a bitch (Middle English biccheSony VAIO PCG-5N2M battery, from Old English bicce, ultimately from Old Norse bikkja). A group of offspring is a litter. The father of a litter is called the sire, and the mother is called the dam. Offspring are, in general, called pups or puppies, from French poupée, until they are about a year old. The process of birth is whelping, from the Old English word hwelp (cf. German Welpe, Dutch welp, Swedish valpa, Icelandic hvelpur) Sony VAIO PCG-3C2M battery.
Taxonomy
In 1753, the father of modern biological taxonomy, Carl Linnaeus, listed among the types of quadrupeds familiar to him, the Latin word for dog, canis. Among the species within this genus, Linnaeus listed the fox, as Canis vulpes, wolves (Canis lupus), and the domestic dog, (Canis canis; see File:Linnaeus - Regnum Animale (1735).png) Sony VAIO PCG-8161M battery.
In later editions, Linnaeus dropped Canis canis and greatly expanded his list of the Canis genus of quadrupeds, and by 1758 included alongside the foxes, wolves, and jackels and many more terms that are now listed as synonyms for domestic dog, including aegyptius (hairless dog), aquaticus, (water dog), and mustelinus (literally “badger dog)” Sony VAIO PCG-8141M battery. Among these were two that later experts have been widely used for domestic dogs as a species: Canis domesticus and, most predominantly, Canis familiaris, the “common” or “familiar” dog.[23]
The domestic dog was accepted as a species in its own right until overwhelming evidence from behavior, vocalizations, morphology, and molecular biology led to the contemporary scientific understanding that a single speciesSony VAIO PCG-3J1M battery, the gray wolf, is the common ancestor for all breeds of domestic dogs. In recognition of this fact, the domestic dog was reclassified in 1993 as Canis lupus familiaris, a subspecies of the gray wolf Canis lupus, by the Smithsonian Institution and the American Society of Mammalogists. C. l. familiaris is listed as the name for the taxon that is broadly used in the scientific community and recommended by ITIS, although Canis familiaris is a recognised synonym. Sony VAIO PCG-3H1M battery
Since that time, C. domesticus and all taxa referring to domestic dogs or subspecies of dog listed by Linnaeus, Johann Friedrich Gmelin in 1792, and Christian Smith in 1839, lost their subspecies status and have been listed as taxonomic synonyms for Canis lupus familiaris
History and evolution
Main articles: Origin of the domestic dog and Gray wolf
Ancient Greek rhyton in the shape of a dog's head, made by Brygos, early 5th century BC. Jérôme Carcopino Museum, Department of Archaeology, AleriaSony VAIO PCG-3F1M battery
Domestic dogs inherited complex behaviors from their wolf ancestors, which would have been pack hunters with complex body language. These sophisticated forms of social cognition and communication may account for their trainability, playfulness, and ability to fit into human households and social situations, and these attributes have given dogs a relationship with humans that has enabled them to become one of the most successful species on the planet today.
Sony VAIO PCG-3C1M battery
Although experts largely disagree over the details of dog domestication, it is agreed that human interaction played a significant role in shaping the subspecies.[29] Domestication may have occurred initially in separate areas particularly Siberia and Europe. Currently it is thought domestication of our current lineage of dog occurred sometime as early as 15,000 years ago and arguably as late as 8500 years agoSony VAIO PCG-9Z2L battery. Shortly after the latest domestication, dogs became ubiquitous in human populations, and spread throughout the world.
Emigrants from Siberia likely crossed the Bering Strait with dogs in their company, and some experts[who?] suggest the use of sled dogs may have been critical to the success of the waves that entered North America roughly 12,000 years ago,[citation needed] although the earliest archaeological evidence of dog-like canids in North America dates from about 9,000 years agoSony VAIO PCG-9Z1L battery.[30] Dogs were an important part of life for the Athabascan population in North America, and were their only domesticated animal. Dogs also carried much of the load in the migration of the Apache and Navajo tribes 1,400 years ago. Use of dogs as pack animals in these cultures often persisted after the introduction of the horse to North America. Sony VAIO PCG-9131L battery
The current consensus among biologists and archaeologists is that the dating of first domestication is indeterminate, although more recent evidence shows isolated domestication events as early as 33,000 years ago. There is conclusive evidence the present lineage of dogs genetically diverged from their wolf ancestors at least 15,000 years agoSony VAIO PCG-8161L battery, but some believe domestication to have occurred earlier.[29] Evidence is accruing that there were previous domestication events, but that those lineages died out.[7]
It is not known whether humans domesticated the wolf as such to initiate dog's divergence from its ancestors, or whether dog's evolutionary path had already taken a different course prior to domesticationSony VAIO PCG-8152L battery. For example, it is hypothesized that some wolves gathered around the campsites of paleolithic camps to scavenge refuse, and associated evolutionary pressure developed that favored those who were less frightened by, and keener in approaching, humans.
Tesem, an old Egyptian sighthound-like dog.
The bulk of the scientific evidence for the evolution of the domestic dog stems from morphological studies of archaeological findings and mitochondrial DNA studiesSony VAIO PCG-8141L battery. The divergence date of roughly 15,000 years ago is based in part on archaeological evidence that demonstrates the domestication of dogs occurred more than 15,000 years ago, and some genetic evidence indicates the domestication of dogs from their wolf ancestors began in the late Upper Paleolithic close to the Pleistocene/Holocene boundary, between 17,000 and 14,000 years ago. Sony VAIO PCG-8131L battery But there is a wide range of other, contradictory findings that make this issue controversial.[citation needed] There are findings beginning currently at 33,000 years ago distinctly placing them as domesticated dogs evidenced not only by shortening of the muzzle but widening as well as crowding of teeth.
Archaeological evidence suggests the latest dogs could have diverged from wolves was roughly 15,000 years ago, although it is possible they diverged much earlier. Sony VAIO PCG-81312L battery In 2008, a team of international scientists released findings from an excavation at Goyet Cave in Belgium declaring a large, toothy canine existed 31,700 years ago and ate a diet of horse, musk ox and reindeer.[37]
Prior to this Belgian discovery, the earliest dog fossils were two large skulls from Russia and a mandible from Germany dated from roughly 14,000 years ago. Sony VAIO PCG-81214L batteryRemains of smaller dogs from Natufian cave deposits in the Middle East, including the earliest burial of a human being with a domestic dog, have been dated to around 10,000 to 12,000 years ago.[5][38] There is a great deal of archaeological evidence for dogs throughout Europe and Asia around this period and through the next two thousand years (roughly 8,000 to 10,000 years ago) Sony VAIO PCG-81115L battery, with fossils uncovered in Germany, the French Alps, and Iraq, and cave paintings in Turkey.[24] The oldest remains of a domesticated dog in the Americas were found in Texas and have been dated to about 9,400 years ago.[39]
DNA studies
DNA studies have provided a wide range of possible divergence dates, from 15,000 to 40,000 years ago,[5] to as much as 100,000 to 140,000 years ago.[40] These results depend on a number of assumptions. Sony VAIO PCG-81114L battery Genetic studies are based on comparisons of genetic diversity between species, and depend on a calibration date. Some estimates of divergence dates from DNA evidence use an estimated wolf-coyote divergence date of roughly 700,000 years ago as a calibration.[41] If this estimate is incorrect, and the actual wolf-coyote divergence is closer to one or two million years ago, or more, then the DNA evidence that supports specific dog-wolf divergence dates would be interpreted very differentlySony VAIO PCG-81113L battery.
Furthermore, it is believed the genetic diversity of wolves has been in decline for the last 200 years, and that the genetic diversity of dogs has been reduced by selective breeding. This could significantly bias DNA analyses to support an earlier divergence date. The genetic evidence for the domestication event occurring in East Asia is also subject to violations of assumptionsSony VAIO PCG-7142L battery. These conclusions are based on the location of maximal genetic divergence, and assume hybridization does not occur, and that breeds remain geographically localized. Although these assumptions hold for many species, there is good reason to believe that they do not hold for canines.[24]
Genetic analyses indicate all dogs are likely descended from a handful of domestication events with a small number of founding females, Sony VAIO PCG-7141L battery although there is evidence domesticated dogs interbred with local populations of wild wolves on several occasions.[5] Data suggest dogs first diverged from wolves in East Asia, and these domesticated dogs then quickly migrated throughout the world, reaching the North American continent around 8000 BC.[5] The oldest groups of dogs, which show the greatest genetic variability and are the most similar to their wolf ancestorsSony VAIO PCG-71111L battery, are primarily Asian and African breeds, including the Basenji, Lhasa Apso, and Siberian Husky.[43] Some breeds thought to be very old, such as the Pharaoh Hound, Ibizan Hound, and Norwegian Elkhound, are now known to have been created more recently.[43]
A great deal of controversy surrounds the evolutionary framework for the domestication of dogs.[24] Although it is widely claimed that "man domesticated the wolf," Sony VAIO PCG-61411L battery man may not have taken such a proactive role in the process.[24] The nature of the interaction between man and wolf that led to domestication is unknown and controversial. At least three early species of the Homo genus began spreading out of Africa roughly 400,000 years ago, and thus lived for a considerable time in contact with canine species. Sony VAIO PCG-61112L battery
Despite this, there is no evidence of any adaptation of canine species to the presence of the close relatives of modern man. If dogs were domesticated, as believed, roughly 15,000 years ago, the event (or events) would have coincided with a large expansion in human territory and the development of agricultureSony VAIO PCG-61111L battery. This has led some biologists to suggest one of the forces that led to the domestication of dogs was a shift in human lifestyle in the form of established human settlements. Permanent settlements would have coincided with a greater amount of disposable food and would have created a barrier between wild and anthropogenic canine populations. Sony VAIO PCG-5T4L battery
Roles with humans
A Siberian Husky used as a pack animal
Early roles
Wolves, and their dog descendants, would have derived significant benefits from living in human camps—more safety, more reliable food, lesser caloric needs, and more chance to breed.[45] They would have benefited from humans’ upright gait that gives them larger range over which to see potential predators and prey, as well as color vision that, at least by day, gives humans better visual discrimination. Sony VAIO PCG-5T3L batteryCamp dogs would also have benefitted from human tool use, as in bringing down larger prey and controlling fire for a range of purposes.[45]
Humans would also have derived enormous benefit from the dogs associated with their camps.[46] For instance, dogs would have improved sanitation by cleaning up food scraps.[46] Dogs may have provided warmth, as referred to in the Australian Aboriginal expression “three dog night” (an exceptionally cold night) Sony VAIO PCG-5T2L battery, and they would have alerted the camp to the presence of predators or strangers, using their acute hearing to provide an early warning.[46]
Anthropologists believe the most significant benefit would have been the use of dogs' sensitive sense of smell to assist with the hunt.[46] The relationship between the presence of a dog and success in the hunt is often mentioned as a primary reason for the domestication of the wolfSony VAIO PCG-5S3L battery, and a 2004 study of hunter groups with and without a dog gives quantitative support to the hypothesis that the benefits of cooperative hunting was an important factor in wolf domestication.[47]
The cohabitation of dogs and humans would have greatly improved the chances of survival for early human groups, and the domestication of dogs may have been one of the key forces that led to human success. Sony VAIO PCG-5S2L battery
A British Bulldog shares a day at the park.
As pets
"The most widespread form of interspecies bonding occurs between humans and dogs"[46] and the keeping of dogs as companions, particularly by elites, has a long history.[49] However, pet dog populations grew significantly after World War II as suburbanization increased.[49] In the 1950s and 1960s, dogs were kept outside more often than they tend to be today Sony VAIO PCG-5S1L battery (using the expression “in the doghouse” to describe exclusion from the group signifies the distance between the doghouse and the home) and were still primarily functional, acting as a guard, children’s playmate, or walking companion. From the 1980s, there have been changes in the role of the pet dog, such as the increased role of dogs in the emotional support of their owners. Sony VAIO PCG-5R2L battery People and dogs have become increasingly integrated and implicated in each other’s lives,[52] to the point where pet dogs actively shape the way a family and home are experienced.[53]
There have been two major trends in the changing status of pet dogs. The first has been the ‘commodification’ of the dog, shaping it to conform to human expectations of personality and behaviour.[53] The second has been the broadening of the concept of the family and the home to include dogs-as-dogs within everyday routines and practices. Sony VAIO PCG-5R1L battery
There are a vast range of commodity forms available to transform a pet dog into an ideal companion.[54] The list of goods, services and places available is enormous: from dog perfumes, couture, furniture and housing, to dog groomers, therapists, trainers and care-takers, dog cafes, spas, parks and beaches, and dog hotels, airlines and cemeteries. Sony VAIO PCG-5P4L battery While dog training as an organized activity can be traced back to the 18th century, in the last decades of the 20th century it became a high profile issue as many normal dog behaviors such as barking, jumping up, digging, rolling in dung, fighting, and urine marking became increasingly incompatible with the new role of a pet dog.[55] Dog training books, classes and television programs proliferated as the process of commodifying the pet dog continued. Sony VAIO PCG-5P2L battery
A pet dog taking part in Christmas traditions
The majority of contemporary dog owners describe their dog as part of the family,[53] although some ambivalence about the relationship is evident in the popular reconceptualization of the dog-human family as a pack.[53] A dominance model of dog-human relationships has been promoted by some dog trainers, such as on the television program Dog WhispererSony VAIO PCG-5N4L battery. However it has been disputed that "trying to achieve status" is characteristic of dog–human interactions.[57] Pet dogs play an active role in family life; for example, a study of conversations in dog-human families showed how family members use the dog as a resource, talking to the dog, or talking through the dog, to mediate their interactions with each other.Sony VAIO PCG-5N2L battery
Another study of dogs’ roles in families showed many dogs have set tasks or routines undertaken as family members, the most common of which was helping with the washing-up by licking the plates in the dishwasher, and bringing in the newspaper from the lawn.[53] Increasingly, human family members are engaging in activities centred on the perceived needs and interests of the dog, or in which the dog is an integral partner, such as Dog Dancing and Doga. Sony VAIO PCG-51513L battery
According to the statistics published by the American Pet Products Manufacturers Association in the National Pet Owner Survey in 2009–2010, it is estimated there are 77.5 million dog owners in the United States.[59] The same survey shows nearly 40% of American households own at least one dog, of which 67% own just one dogSony VAIO PCG-51511L battery, 25% two dogs and nearly 9% more than two dogs. There does not seem to be any gender preference among dogs as pets, as the statistical data reveal an equal number of female and male dog pets. Yet, although several programs are undergoing to promote pet adoption, less than a fifth of the owned dogs come from a shelterSony VAIO PCG-51412L battery.
Work
Dogs have lived and worked with humans in so many roles that they have earned the unique nickname, "man's best friend",[60] a phrase used in other languages as well. They have been bred for herding livestock,[61] hunting (e.g. pointers and hounds),[62] rodent control,[3] guarding, helping fishermen with nets, detection dogs, and pulling loads, in addition to their roles as companionsSony VAIO PCG-51411L battery.
Book of the Hunt, Gaston III, Count of Foix, 1387–88.
Service dogs such as guide dogs, utility dogs, assistance dogs, hearing dogs, and psychological therapy dogs provide assistance to individuals with physical or mental disabilities.[63][64] Some dogs owned by epileptics have been shown to alert their handler when the handler shows signs of an impending seizure, sometimes well in advance of onset, allowing the owner to seek safety, medication, or medical care. Sony VAIO PCG-51312L battery
Dogs included in human activities in terms of helping out humans are usually called working dogs. Dogs of several breeds are considered working dogs. Some working dog breeds include Akita, Alaskan Malamute, Anatolian Shepherd Dog, Bernese Mountain Dog, Black Russian Terrier, Boxer, Bullmastiff, Doberman Pinscher, Dogue de Bordeaux, German Pinscher, German Shepherd, Sony VAIO PCG-51311L batteryGiant Schnauzer, Great Dane, Great Pyrenees, Great Swiss Mountain Dog, Komondor, Kuvasz, Mastiff, Neapolitan Mastiff, Newfoundland, Portuguese Water Dog, Rottweiler, Saint Bernard, Samoyed, Siberian Husky, Standard Schnauzer, and Tibetan Mastiff.
Sports and shows
See also: Conformation show
Owners of dogs often enter them in competitions[67] such as breed conformation shows or sports, including racing and sledding.
In conformation shows, also referred to as breed shows, a judge familiar with the specific dog breed evaluates individual purebred dogs for conformity with their established breed type as described in the breed standardSony VAIO PCG-51211L battery. As the breed standard only deals with the externally observable qualities of the dog (such as appearance, movement, and temperament), separately tested qualities (such as ability or health) are not part of the judging in conformation shows.
As a food source
See also: Dog meat
Fried dog meat offered as speciality, among lamb and fish. Shaoyang, Hunan Province, China
Dog meat is consumed in some East Asian countries, including Korea, China, and Vietnam, a practice that dates back to antiquity. Sony VAIO PCG-41112L battery It is estimated that 13–16 million dogs are killed and consumed in Asia every year.[69] The BBC claims that, in 1999, more than 6,000 restaurants served soups made from dog meat in South Korea.[70] In Korea, the primary dog breed raised for meat, the nureongi (누렁이), differs from those breeds raised for pets that Koreans may keep in their homes. Sony VAIO PCG-3A4L battery
The most popular Korean dog dish is gaejang-guk (also called bosintang), a spicy stew meant to balance the body's heat during the summer months; followers of the custom claim this is done to ensure good health by balancing one's gi, or vital energy of the body. A 19th century version of gaejang-guk explains that the dish is prepared by boiling dog meat with scallions and chili powderSony VAIO PCG-3A3L battery. Variations of the dish contain chicken and bamboo shoots. While the dishes are still popular in Korea with a segment of the population, dog is not as widely consumed as beef, chicken, and pork.
Other cultures, such as Polynesia and pre-Columbian Mexico, also consumed dog meat in their history. However, Western, South Asian, African, and Middle Eastern cultures, in general, regard consumption of dog meat as tabooSony VAIO PCG-3A2L battery. In some places, however, such as in rural areas of Poland, dog fat is believed to have medicinal properties—being good for the lungs for instance.[73]
A CNN report in China dated March 2010 interviews a dog meat vendor who states that most of the dogs that are available for selling to restaurant are raised in special farms but that there is always a chance that a sold dog is someone's lost pet, although dog pet breeds are not considered edible. Sony VAIO PCG-3A1L battery
Health risks to humans
Main article: Dog attack
It is estimated that two percent of the US population, 4.7 million people, are bitten each year.[75] In the 1980s and 1990s the US averaged 17 fatalities per year, while in the 2000s this has increased to 26. 77% of dog bites are from the pet of family or friends, and 50% of attacks occur on the dog owner's property. Sony VAIO PCG-394L battery
A Colorado study found bites in children were less severe than bites in adults.[77] The incidence of dog bites in the U.S. is 12.9 per 10,000 inhabitants, but for boys aged 5 to 9, the incidence rate is 60.7 per 10,000. Moreover, children have a much higher chance to be bitten in the face or neck.[78] Sharp claws with powerful muscles behind them can lacerate flesh in a scratch that can lead to serious infections. Sony VAIO PCG-393L battery
In the UK between 2003 and 2004, there were 5,868 dog attacks on humans, resulting in 5,770 working days lost in sick leave.[80]
In the United States, cats and dogs are a factor in more than 86,000 falls each year.[81] It has been estimated around 2% of dog-related injuries treated in UK hospitals are domestic accidents. The same study found that while dog involvement in road traffic accidents was difficult to quantify, dog-associated road accidents involving injury more commonly involved two-wheeled vehicles. Sony VAIO PCG-391L battery
Toxocara canis (dog roundworm) eggs in dog feces can cause toxocariasis. In the United States, about 10,000 cases of Toxocara infection are reported in humans each year, and almost 14% of the U.S. population is infected.[83] In Great Britain, 24% of soil samples taken from public parks contained T. canis eggs.[84] Untreated toxocariasis can cause retinal damage and decreased vision. Sony VAIO PCG-384L battery Dog feces can also contain hookworms that cause cutaneous larva migrans in humans.
Health benefits for humans
A human cuddles a Doberman puppy.
A growing body of research indicates the companionship of a dog can enhance human physical health and psychological wellbeing.[89] Dog and cat owners have been shown to have better mental and physical health than nonowners, making fewer visits to the doctor and being less likely to be on medication than nonowners. Sony VAIO PCG-383L battery In one study, new pet owners reported a highly significant reduction in minor health problems during the first month following pet acquisition, and this effect was sustained in dog owners through to the end of the study.[91]
In addition, dog owners took considerably more physical exercise than cat owners and people without pets. The group without pets exhibited no statistically significant changes in health or behaviourSony VAIO PCG-382L battery. The results provide evidence that pet acquisition may have positive effects on human health and behaviour, and that for dog owners these effects are relatively long term.[91] Pet ownership has also been associated with increased coronary artery disease survival, with dog owners being significantly less likely to die within one year of an acute myocardial infarction than those who did not own dogs.Sony VAIO PCG-381L battery
Gunnar Kaasen and Balto, the lead dog on the last relay team of the 1925 serum run to Nome.
The health benefits of dogs can result from contact with dogs, not just from dog ownership. For example, when in the presence of a pet dog, people show reductions in cardiovascular, behavioral, and psychological indicators of anxiety. Sony VAIO PCG-7185L battery The benefits of contact with a dog also include social support, as dogs are able to not only provide companionship and social support themselves, but also to act as facilitators of social interactions between humans.[94] One study indicated that wheelchair users experience more positive social interactions with strangers when they are accompanied by a dog than when they are not. Sony VAIO PCG-7184L battery
The practice of using dogs and other animals as a part of therapy dates back to the late 18th century, when animals were introduced into mental institutions to help socialize patients with mental disorders.[96] Animal-assisted intervention research has shown that animal-assisted therapy with a dog can increase a person with Alzheimer’s disease’s social behavioursSony VAIO PCG-7183L battery, such as smiling and laughing.[97] One study demonstrated that children with ADHD and conduct disorders who participated in an education program with dogs and other animals showed increased attendance, increased knowledge and skill objectives, and decreased antisocial and violent behavior compared to those who were not in an animal-assisted program. Sony VAIO PCG-7182L battery
Shelters
Main article: Animal shelter
Every year, between 6 and 8 million dogs and cats enter US animal shelters.[99] The Humane Society of the United States (HSUS) estimates that approximately 3 to 4 million dogs and cats are euthanized yearly in shelters across the United States. However, the percentage of dogs in US animal shelters that are eventually adopted and removed from the sheltersSony VAIO PCG-7181L battery by their new owners has increased since the mid 1990s from around 25% up to around 60–75% in the mid first decade of the 21st century.[101]
Pets entering the shelters are euthanized in countries all over the world because of the lack of financial provisions to take care of these animals. Most shelters complain of not having enough resources to feed the pets and by being constrained to kill them, as the likelihood for all of them to find an owner is very small. In poor countries, euthanasia is usually violentSony VAIO PCG-7174L battery.
Biology
Main article: Dog anatomy
Lateral view of a dog's bone structure.
Domestic dogs have been selectively bred for millennia for various behaviors, sensory capabilities, and physical attributes.[3] Modern dog breeds show more variation in size, appearance, and behavior than any other domestic animal. Nevertheless, their morphology is based on that of their wild ancestors, gray wolves.[3] Dogs are predators and scavengersSony VAIO PCG-7173L battery, and like many other predatory mammals, the dog has powerful muscles, fused wrist bones, a cardiovascular system that supports both sprinting and endurance, and teeth for catching and tearing.
Dogs are highly variable in height and weight. The smallest known adult dog was a Yorkshire Terrier, that stood only 6.3 centimetres (2.5 in) at the shoulder, 9.5 cm (3.7 in) in length along the head-and-body, and weighed only 113 grams (4.0 oz)Sony VAIO PCG-7172L battery. The largest known dog was an English Mastiff which weighed 155.6 kilograms (343 lb) and was 250 cm (98 in) from the snout to the tail.[102] The tallest dog is a Great Dane that stands 106.7 cm (42.0 in) at the shoulder.
Senses
Vision
Dog's visual colour perception compared with humans.
Like most mammals, dogs are dichromats and have color vision equivalent to red-green color blindness in humans (deuteranopia). Dogs are less sensitive to differences in grey shades than humans and also can detect brightness at about half the accuracy of humans. Sony VAIO PCG-7171L battery
The dog's visual system has evolved to aid proficient hunting.[104] While a dog's visual acuity is poor (that of a poodle's has been estimated to translate to a Snellen rating of 20/75[104]), their visual discrimination for moving objects is very high; dogs have been shown to be able to discriminate between humans (e.g., identifying their owner) at a range of between 800 and 900 m, however this range decreases to 500–600 m if the object is stationary. Sony VAIO PCG-7162L battery
Dogs have a temporal resolution of between 60 and 70 Hz, which explains why many dogs struggle to watch television, as most such modern screens are optimized for humans at 50–60 Hz.[108] Dogs can detect a change in movement that exists in a single diopter of space within their eye. Humans, by comparison, require a change of between 10 and 20 diopters to detect movementSony VAIO PCG-7161L battery.
The physiology of a dog ear.
As crepuscular hunters, dogs often rely on their vision in low light situations: They have very large pupils, a high density of rods in the fovea, an increased flicker rate, and a tapetum lucidum.[104] The tapetum is a reflective surface behind the retina that reflects light to give the photoreceptors a second chance to catch the photons. There is also a relationship between body size and overall diameter of the eyeSony VAIO PCG-7154L battery. A range of 9.5 and 11.6 mm can be found between various breeds of dogs. This 20% variance can be substantial and is associated as an adaptation toward superior night vision.[111]
The eyes of different breeds of dogs have different shapes, dimensions, and retina configurations.[112] Many long-nosed breeds have a "visual streak" – a wide foveal region that runs across the width of the retina and gives them a very wide field of excellent visionSony VAIO PCG-7153L battery. Some long-muzzled breeds, in particular, the sighthounds, have a field of vision up to 270° (compared to 180° for humans). Short-nosed breeds, on the other hand, have an "area centralis": a central patch with up to three times the density of nerve endings as the visual streak, giving them detailed sight much more like a human's. Some broad-headed breeds with short noses have a field of vision similar to that of humans. Sony VAIO PCG-7152L battery
Most breeds have good vision, but some show a genetic predisposition for myopia – such as Rottweilers, with which one out of every two has been found to be myopic.[104] Dogs also have a greater divergence of the eye axis than humans, enabling them to rotate their pupils farther in any direction. The divergence of the eye axis of dogs ranges from 12–25° depending on the breed.Sony VAIO PCG-7151L battery
Experimentation has proven that dogs can distinguish between complex visual images such as that of a cube or a prism. Dogs also show attraction to static visual images such as the silhouette of a dog on a screen, their own reflections, or videos of dogs; however, their interest declines sharply once they are unable to make social contact with the image.
Sony VAIO PCG-7148L battery
Hearing
The frequency range of dog hearing is approximately 40 Hz to 60,000 Hz,[114] which means that dogs can detect sounds far beyond the upper limit of the human auditory spectrum. In addition, dogs have ear mobility, which allows them to rapidly pinpoint the exact location of a sound. Eighteen or more muscles can tilt, rotate, raise, or lower a dog's earSony VPCW21M2E/WI battery. A dog can identify a sound's location much faster than a human can, as well as hear sounds at four times the distance.[116]
Smell
The wet, textured nose of a dog
While the human brain is dominated by a large visual cortex, the dog brain is dominated by an olfactory cortex.[104] The olfactory bulb in dogs is roughly forty times bigger than the olfactory bulb in humans, relative to total brain size, with 125 to 220 million smell-sensitive receptors.[104] The bloodhound exceeds this standard with nearly 300 million receptors. Sony VPCW21C7E battery
Consequently, it has been estimated that dogs, in general, have an olfactory sense ranging from one hundred thousand to one million times more sensitive than a human's. In some dog breeds, such as bloodhounds, the olfactory sense may be up to 100 million times greater than a human's.[117] The wet nose is essential for determining the direction of the air current containing the smellSony VPCW12S1E/WZ battery. Cold receptors in the skin are sensitive to the cooling of the skin by evaporation of the moisture by air currents.[118]
Physical characteristics
Main article: Dog anatomy
Coat
Main article: Coat (dog)
A heavy winter coat with countershading in a mixed-breed dog
The coats of domestic dogs are of two varieties: "double" being common with dogs (as well as wolves) originating from colder climates, made up of a coarse guard hair and a soft down hair, or "single", with the topcoat onlySony VPCW12S1E/W battery.
Domestic dogs often display the remnants of countershading, a common natural camouflage pattern. A countershaded animal will have dark coloring on its upper surfaces and light coloring below,[119] which reduces its general visibility. Thus, many breeds will have an occasional "blaze", stripe, or "star" of white fur on their chest or underside. Sony VPCW12S1E/T battery
Tail
See also: Docking
There are many different shapes for dog tails: straight, straight up, sickle, curled, or cork-screw. As with many canids, one of the primary functions of a dog's tail is to communicate their emotional state, which can be important in getting along with others. In some hunting dogs, however, the tail is traditionally docked to avoid injuries.[121] In some breeds, puppies can be born with a short tail or no tail at allSony VPCW12S1E/P battery.
Types and breeds
Main article: Dog breed
Further information: Dog type
Cavalier King Charles Spaniels demonstrate with-breed variation.
While all dogs are genetically very similar,[5] natural selection and selective breeding have reinforced certain characteristics in certain populations of dogs, giving rise to dog types and dog breeds. Dog types are broad categories based on function, genetics, or characteristics.[123] Dog breeds are groups of animals that possess a set of inherited characteristics that distinguishes them from other animals within the same speciesSony VPCW11S1E/W battery. Modern dog breeds are non-scientific classifications of dogs kept by modern kennel clubs.
Purebred dogs of one breed are genetically distinguishable from purebred dogs of other breeds,[43] but the means by which kennel clubs classify dogs is unsystematic. Systematic analyses of the dog genome has revealed only four major types of dogs that can be said to be statistically distinct.[43] These include the "old world dogs" (e.g., Malamute and Shar Pei) Sony VPCW11S1E/T battery, "Mastiff"-type (e.g., English Mastiff), "herding"-type (e.g., Border Collie), and "all others" (also called "modern"- or "hunting"-type).
Health
Main article: Dog health
Further information: Category:Dog health
See also: CVBD
Dogs are susceptible to various diseases, ailments, and poisons, some of which can affect humans. To defend against many common diseases, dogs are often vaccinated.
A mixed-breed dog
Some breeds of dogs are prone to certain genetic ailments such as elbow or hip dysplasia, blindness, deafness, pulmonic stenosis, cleft palate, and trick knees. Two serious medical conditions particularly affecting dogs are pyometra, affecting unspayed females of all types and agesSony VPCW11S1E/P battery, and bloat, which affects the larger breeds or deep-chested dogs. Both of these are acute conditions, and can kill rapidly. Dogs are also susceptible to parasites such as fleas, ticks, and mites, as well as hookworm, tapeworm, roundworm, and heartworm.
Dogs are highly susceptible to theobromine poisoning, typically from ingestion of chocolate. Theobromine is toxic to dogs because, although the dog's metabolism is capable of breaking down the chemicalSony VPCYA1S1E/B battery, the process is so slow that even small amounts of chocolate can be fatal, especially dark chocolate.
Dogs are also vulnerable to some of the same health conditions as humans, including diabetes, dental and heart disease, epilepsy, cancer, hypothyroidism, and arthritis.[125]
Mortality
Main article: Aging in dogs
The typical lifespan of dogs varies widely among breeds, but for most the median longevity, the age at which half the dogs in a population have died and half are still alive, ranges from 10 to 13 years. Individual dogs may live well beyond the median of their breedSony VPCYA1V9E/B battery.
The breed with the shortest lifespan (among breeds for which there is a questionnaire survey with a reasonable sample size) is the Dogue de Bordeaux, with a median longevity of about 5.2 years, but several breeds, including Miniature Bull Terriers, Bloodhounds, and Irish Wolfhounds are nearly as short-lived, with median longevities of 6 to 7 years. Sony VPCY21S1E/SI battery
The longest-lived breeds, including Toy Poodles, Japanese Spitz, Border Terriers, and Tibetan Spaniels, have median longevities of 14 to 15 years.[129] The median longevity of mixed-breed dogs, taken as an average of all sizes, is one or more years longer than that of purebred dogs when all breeds are averaged. The dog widely reported to be the longest-lived is "Bluey," Sony VPCY21S1E/L batterywho died in 1939 and was claimed to be 29.5 years old at the time of his death; however, the Bluey record is anecdotal and unverified.[131] On December 5, 2011, Pusuke, the world's oldest living dog recognized by Guinness Book of World Records, died aged 26 years and 9 months.[132]
Predation
Although wild dogs, like wolves, are apex predators, they can be killed in territory disputes with wild animals. Sony VPCY21S1E/G batteryFurthermore, in areas where both dogs and other large predators live, dogs can be a major food source for big cats or canines. Reports from Croatia indicate wolves kill dogs more frequently than they kill sheep. Wolves in Russia apparently limit feral dog populations. In Wisconsin, more compensation has been paid for dog losses than livestock.[133] Some wolf pairs have been reported to prey on dogs by having one wolf lure the dog out into heavy brush where the second animal waits in ambush. Sony VPCY11S1E/S battery In some instances, wolves have displayed an uncharacteristic fearlessness of humans and buildings when attacking dogs, to the extent that they[which?] have to be beaten off or killed.[135]
Coyotes and big cats have also been known to attack dogs. Leopards in particular are known to have a predilection for dogs, and have been recorded to kill and consume them regardless of the dog's size or ferocity.[136] Tigers in Manchuria, Indochina, Indonesia, and Malaysia are reputed to kill dogs with the same vigor as leopards. Sony VPCY11S1E battery Striped Hyenas are major predators of village dogs in Turkmenistan, India, and the Caucasus.[138] Reptiles such as alligators and pythons have been known to kill and eat dogs.
Diet
Dog food
Golden Retriever gnawing a pig's foot
Despite their descent from wolves and classification as Carnivora, dogs are variously described in scholarly and other writings as carnivores or omnivores. Unlike obligate carnivores, such as the cat family with its shorter small intestineSony VPCZ11X9E/B battery, dogs can adapt to a wide-ranging diet, and are not dependent on meat-specific protein nor a very high level of protein in order to fulfill their basic dietary requirements. Dogs will healthily digest a variety of foods, including vegetables and grains, and can consume a large proportion of these in their diet. Sony VPCZ11Z9E/B battery
A number of common human foods and household ingestibles are toxic to dogs, including chocolate solids (theobromine poisoning), onion and garlic (thiosulphate, sulfoxide or disulfide poisoning),[144] grapes and raisins, macadamia nuts, as well as various plants and other potentially ingested materialsSony VPCZ12M9E/B battery.
Reproduction
Two dogs copulating on a beach
Main article: Canine reproduction
In domestic dogs, sexual maturity begins to happen around age six to twelve months for both males and females,[3][147] although this can be delayed until up to two years old for some large breeds. This is the time at which female dogs will have their first estrous cycle. They will experience subsequent estrous cycles biannually, during which the body prepares for pregnancySony VPCZ12X9E/X battery. At the peak of the cycle, females will come into estrus, being mentally and physically receptive to copulation.[3] Because the ova survive and are capable of being fertilized for a week after ovulation, it is possible for a female to mate with more than one male.[3]
Dogs bear their litters roughly 56 to 72 days after fertilization,[3][148] with an average of 63 days, although the length of gestation can vary. An average litter consists of about six puppies, Sony VPCZ12Z9E/X battery though this number may vary widely based on the breed of dog. In general, toy dogs produce from one to four puppies in each litter, while much larger breeds may average as many as twelve.
Some dog breeds have acquired traits through selective breeding that interfere with reproduction. Male French Bulldogs, for instance, are incapable of mounting the female. For many dogs of this breed, the female must be artificially inseminated in order to reproduce. Sony VPCZ138GA battery
Neutering
The examples and perspective in this article deal primarily with the United States and do not represent a worldwide view of the subject. Please improve this article and discuss the issue on the talk page. (March 2012)
A feral dog from Sri Lanka nursing her four puppies
Neutering refers to the sterilization of animals, usually by removal of the male's testicles or the female's ovaries and uterus, in order to eliminate the ability to procreate and reduce sex driveSony VPCZ13M9E/B battery. Because of the overpopulation of dogs in some countries, many animal control agencies, such as the American Society for the Prevention of Cruelty to Animals (ASPCA), advise that dogs not intended for further breeding should be neutered, so that they do not have undesired puppies that may have to later be euthanized.[151]
According to the Humane Society of the United States, 3–4 million dogs and cats are put down each year in the United States and many more are confined to cages in shelters because there are many more animals than there are homesSony VPCZ13M9E/X battery. Spaying or castrating dogs helps keep overpopulation down.[152] Local humane societies, SPCAs, and other animal protection organizations urge people to neuter their pets and to adopt animals from shelters instead of purchasing them.
Neutering reduces problems caused by hypersexuality, especially in male dogs.[153] Spayed female dogs are less likely to develop some forms of cancer, affecting mammary glands, ovaries, and other reproductive organs. Sony VPCZ13V9E batteryHowever, neutering increases the risk of urinary incontinence in female dogs,[155] and prostate cancer in males,[156] as well as osteosarcoma, hemangiosarcoma, cruciate ligament rupture, obesity, and diabetes mellitus in either gender.[157]
Intelligence and behavior
Intelligence
Main article: Dog intelligence
The Border Collie is considered to be one of the most intelligent breeds.
The domestic dog has a predisposition to exhibit a social intelligence that is uncommon in the animal world.[104] Dogs are capable of learning in a number of ways, such as through simple reinforcement (e.g., classical or operant conditioning) and by observation. Sony VPCZ13V9E/X battery
Dogs go through a series of stages of cognitive development. As with humans, the understanding that objects not being actively perceived still remain in existence (called object permanence) is not present at birth. It develops as the young dog learns to interact intentionally with objects around it, at roughly 8 weeks of age. Sony VPCZ13Z9E/X battery
Puppies learn behaviors quickly by following examples set by experienced dogs.[104] This form of intelligence is not peculiar to those tasks dogs have been bred to perform, but can be generalized to myriad abstract problems. For example, Dachshund puppies that watched an experienced dog pull a cart by tugging on an attached piece of ribbon in order to get a reward from inside the cart learned the task fifteen times faster than those left to solve the problem on their own. Sony VPCZ21M9E battery
Dogs can also learn by mimicking human behaviors. In one study, puppies were presented with a box, and shown that, when a handler pressed a lever, a ball would roll out of the box. The handler then allowed the puppy to play with the ball, making it an intrinsic reward. The pups were then allowed to interact with the box. Roughly three-quarters of the puppies subsequently touched the leverSony VPCZ21Q9E battery, and over half successfully released the ball, compared to only 6% in a control group that did not watch the human manipulate the lever.[159] Another study found that handing an object between experimenters who then used the object's name in a sentence successfully taught an observing dog each object's name, allowing the dog to subsequently retrieve the item. Sony VPCZ21V9E battery
Sergeant Stubby wearing his uniform and medals. Stubby participated in four offensives and 17 battles.
Dogs also demonstrate sophisticated social cognition by associating behavioral cues with abstract meanings.[104] One such class of social cognition involves the understanding that others are conscious agents. Research has shown that dogs are capable of interpreting subtle social cuesSony VPCEH3T9E battery, and appear to recognize when a human or dog's attention is focused on them. To test this, researchers devised a task in which a reward was hidden underneath one of two buckets. The experimenter then attempted to communicate with the dog to indicate the location of the reward by using a wide range of signals: tapping the bucket, pointing to the bucket, nodding to the bucket, or simply looking at the bucket. Sony VPCEH3N6E battery The results showed that domestic dogs were better than chimpanzees, wolves, and human infants at this task, and even young puppies with limited exposure to humans performed well.[104]
Psychology research has shown that humans´ gaze instinctively moves to the left in order to watch the right side of a person's face, which is related to use of right hemisphere brain for facial recognition, including human facial emotions. Research at the University of Lincoln (2008) shows that dogs share this instinct when meeting a human beingSony VPCEH3N1E battery, and only when meeting a human being (i.e., not other animals or other dogs). As such they are the only non-primate species known to do so.[162][163]
Stanley Coren, an expert on dog psychology, states that these results demonstrated the social cognition of dogs can exceed that of even our closest genetic relatives, and that this capacity is a recent genetic acquisition that distinguishes the dog from its ancestor, the wolf. Sony VPCEH3D0E battery Studies have also investigated whether dogs engaged in partnered play change their behavior depending on the attention-state of their partner.[164] Those studies showed that play signals were only sent when the dog was holding the attention of its partner. If the partner was distracted, the dog instead engaged in attention-getting behavior before sending a play signal. Sony VPCEH3B1E battery
Coren has also argued that dogs demonstrate a sophisticated theory of mind by engaging in deception, which he supports with a number of anecdotes, including one example wherein a dog hid a stolen treat by sitting on it until the rightful owner of the treat left the room.[104] Although this could have been accidental, Coren suggests that the thief understood that the treat's owner would be unable to find the treat if it were out of viewSony VPCEH2Z1E battery. Together, the empirical data and anecdotal evidence points to dogs possessing at least a limited form of theory of mind.[104][164]
A study found a third of dogs suffered from anxiety when separated from others.[165]
A Border Collie named Chaser has learned the names for 1,022 toys after three years of training, so many that her trainers have had to mark the names of the objects lest they forget themselves. This is higher than Rico, another border collie who could remember at least 200 objects. Sony VPCEH2S9E battery
Behavior
Main article: Dog behavior
Further information: Category:Dog training and behavior
Although dogs have been the subject of a great deal of behaviorist psychology (e.g. Pavlov's dog), they do not enter the world with a psychological "blank slate".[104] Rather, dog behavior is affected by genetic factors as well as environmental factors.[104] Domestic dogs exhibit a number of behaviors and predispositions that were inherited from wolves. Sony VPCEH2Q1E battery
The Gray Wolf is a social animal that has evolved a sophisticated means of communication and social structure. The domestic dog has inherited some of these predispositions, but many of the salient characteristics in dog behavior have been largely shaped by selective breeding by humans. Thus some of these characteristics, such as the dog's highly developed social cognition, are found only in primitive forms in grey wolves. Sony VPCEH2P0E battery
The existence and nature of personality traits in dogs have been studied (15329 dogs of 164 different breeds) and five consistent and stable "narrow traits" identified, described as playfulness, curiosity/fearlessness, chase-proneness, sociability and aggressiveness. A further higher order axis for shyness–boldness was also identified. Sony VPCEH2N1E battery
Sleep
Further information: Sleep (non-human)
The average sleep time of a dog is said to be 10.1 hours per day.[169] Like humans, dogs have two main types of sleep: Slow-wave sleep, then Rapid eye movement sleep sleep, the state in which dreams occur.[170]
Dog growl
A new study in Budapest, Hungary has found that dogs are able to tell how big another dog is just by listening to its growl. A specific growl is used by dogs to protect their food. The research also shows that dogs do not lie about their sizeSony VPCEH2M9E battery, and this is the first time research has shown animals can determine another’s size by the sound it makes. The test, using images of many kinds of dogs, showed a small and big dog and played a growl. The result, showed that 20 of the 24 test dogs looked at the image of the appropriate-sized dog first and looked at it longest. Sony VPCEH2M1E battery
Differences from wolves
Some dogs, like this Tamaskan, look very much like wolves.
Physical characteristics
Further information: Wolf
Compared to equally sized wolves, dogs tend to have 20% smaller skulls, 30% smaller brains,[172] as well as proportionately smaller teeth than other canid species.[173] Dogs require fewer calories to function than wolves. It is thought by certain experts that the dog's limp ears are a result of atrophy of the jaw muscles.[173] The skin of domestic dogs tends to be thicker than that of wolvesSony VPCEH2L9E battery, with some Inuit tribes favoring the former for use as clothing due to its greater resistance to wear and tear in harsh weather.[173]
Behavior
Dogs tend to be poorer than wolves at observational learning, being more responsive to instrumental conditioning.[173] Feral dogs show little of the complex social structure or dominance hierarchy present in wolf packs. For example, unlike wolves, the dominant alpha pairs of a feral dog pack do not force the other members to wait for their turn on a meal when scavenging off a dead ungulate as the whole family is free to join inSony VPCEH2J1E battery. For dogs, other members of their kind are of no help in locating food items, and are more like competitors.[173]
Feral dogs are primarily scavengers, with studies showing that unlike their wild cousins, they are poor ungulate hunters, having little impact on wildlife populations where they are sympatric. However, feral dogs have been reported to be effective hunters of reptiles in the Galápagos Islands, Sony VPCEH2H1E battery and free ranging pet dogs are more prone to predatory behavior toward wild animals.
Domestic dogs can be monogamous.[175] Breeding in feral packs can be, but does not have to be restricted to a dominant alpha pair (such things also occur in wolf packs).[176] Male dogs are unusual among canids by the fact that they mostly seem to play no role in raising their puppies, and do not kill the young of other females to increase their own reproductive success. Sony VPCEH2F1E battery Some sources say that dogs differ from wolves and most other large canid species by the fact that they do not regurgitate food for their young, nor the young of other dogs in the same territory.[173]
An Australian Shepherd-Beagle mix displaying mastery of the "sit" command
However, this difference was not observed in all domestic dogs. Regurgitating of food by the females for the young as well as care for the young by the males has been observed in domestic dogsSony VPCEH2E0E battery, dingos as well as in other feral or semi-feral dogs. Regurgitating of food by the females and direct choosing of only one mate has been observed even in those semi-feral dogs of direct domestic dog ancestry. Also regurgitating of food by males has been observed in free-ranging domestic dogs.
Trainability
Dogs display much greater tractability than tame wolves, and are, in general, much more responsive to coercive techniques involving fearSony VPCEH2D0E battery, aversive stimuli, and force than wolves, which are most responsive toward positive conditioning and rewards.[178] Unlike tame wolves, dogs tend to respond more to voice than hand signals.[179]
Mythology
In mythology, dogs often serve as pets or as watchdogs.[180]
In Greek mythology, Cerberus is a three-headed watchdog who guards the gates of Hades.[180] In Norse mythology, a bloody, four-eyed dog called Garmr guards Helheim.[180] In Persian mythology, two four-eyed dogs guard the Chinvat Bridge. Sony VPCEH2C0E batteryIn Philippine mythology, Kimat who is the pet of Tadaklan, god of thunder, is responsible for lightning. In Welsh mythology, Annwn is guarded by Cŵn Annwn[180]
In Judaism and Islam, dogs are viewed as unclean scavengers.[180] In Christianity, dogs represent faithfulness.[180] In Asian countries such as China, Korea, and Japan, dogs are viewed as kind protectorsSony VPCEH1Z1E battery.
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